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Squeeze…

[T]he testicles are exquisitely sensitive to touch and there is a huge release of adrenalin when there is excessive force applied to these organs. … A heart attack could certainly result from severe testicular pain from squeezing.

Dr. Irwin Goldstein, quoted in Can You Die If Someone Squeezes Your Testicles Hard Enough? [gizmodo.com] by Jesus Diaz